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GUANTANAMO BAY, Cuba (CNN) – Osama bin Laden’s former driver knew the target of one of the hijacked planes on September 11, 2001, prosecutors said as the military commission trial of Salim Hamdan began Tuesday.

Hamdan told U.S. interrogators the fourth plane was aimed at “the dome,” apparently a reference to the U.S. Capitol, the prosecutors said. The plane crashed in a field in Pennsylvania after the passengers fought the hijackers.

But the defense said Hamdan was a low-level driver who knew little about the workings of bin Laden’s al Qaeda network. They said he worked for wages, not to wage attacks against America.

Captured at a roadblock in Afghanistan in November 2001, Hamdan pleaded not guilty at the start of the trial, which will be closely watched as the first full test of the Pentagon’s system for prosecuting alleged terrorists.

He faces a maximum life sentence if convicted of conspiracy and aiding terrorism.

The first prosecution witness, a U.S. soldier who was present when Hamdan was captured, originally said Hamdan was driving a car that had SA-7 surface-to-air missiles intended for al Qaeda. But on cross-examination, the soldier, identified only as Maj. Smith, said he could not be sure Hamdan was the driver of one of the three vehicles carrying the missiles.

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